Nobel laureate Har Gobind Khorana dies at 89

Khorana, 89, who was MIT's Alfred P Sloan Professor of Biology and Chemistry emeritus, died earlier this week.

He won the Nobel Prize in 1968, sharing it with two others, for unravelling the nucleotide sequence of RNA and deciphering the genetic code. He was then with the University of Wisconsin (UW).

He is survived by his daughter, Julia, and son, Dave.

Born in 1922, in a small village called Raipur in Punjab, which is now in Pakistan, Khorana is known as a scientist who revolutionised biochemistry with his pioneering work in DNA chemistry.

"The work that he did in Wisconsin from 1960 to 1970 continues to propel new scientific discoveries and major advances," said Aseem Ansari, professor of biochemistry at the University of Wisconsin-Madison where Khorana taught and did research from 1960 to 1970 before moving to the Massachusetts Institute of Technology (MIT).

It was at Wisconsin that Khorana along with his colleagues worked out mechanisms of RNA codes for the synthesis of proteins, which won him the Nobel Prize.

He shared the prize with Robert Holley of Cornell University and Marshall Nirenberg of the National Institutes of Health.

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